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Physalis heterophylla

Common name:	Clammy Ground Cherry	Family:	Solanaceae
Author:	Nees.	Botanical references:	170, 200, 235
Synonyms:	Physalis ambigua ((Gray) Britton.)
Known Hazards:	All parts of the plant, except the fruit, are poisonous[19, 65].
Range:	N. America - New Brunswick to Saskatchewan, Florida, Colorado and Texas.
Habitat:	Dry open woods[43]. Rich soils, especially where the surface has been disturbed[235].
Edibility Rating (1-5):	3	Medicinal Rating (1-5):	1

Other Possible Synonyms:	From various places across the web, may not be correct. See below.
	P. heterophylla var. ambigua[B,P] P. heterophylla var. clavipes[B,P] P. heterophylla var. heterophylla[B,P] P. heterophylla var. nyctaginea[B,P] P. heterophylla var. villosa[B,P] P. nyctaginea[B,P] P. sinuata[B,P]
Other Common Names:	From various places around the Web, may not be correct. See below.
	Clammy Ground-cherry [B], Clammy Groundcherry [P],
Epithets:	From a Dictionary of Botanical Epithets
	heterophylla = differently leaved
Systematics:	From a USDA Plants Database
	Order: Solanales. Potato family
Noxious, Invasive and Injurious Weeds	From USDA PLANTS database, Weeds Australia , DEFRA Injurious Weeds
	Listed as noxious/invasive for: USA Invasive.

Physical Characteristics

Perennial growing to 0.9m by 1m . It is hardy to zone 8. It is in flower from July to September. The flowers are hermaphrodite (have both male and female organs) and are pollinated by Insects. We rate it 3/5 for edibility and 1/5 for medicinal use.

The plant prefers light (sandy), medium (loamy) and heavy (clay) soils and requires well-drained soil. The plant prefers acid, neutral and basic (alkaline) soils. It can grow in semi-shade (light woodland) or no shade. It requires dry or moist soil.

Habitats and Possible Locations

Cultivated Beds.

Edible Uses

Fruit.

Fruit - raw or cooked[46, 105, 161]. Quite nice raw though rather small[K], the fruit can also be made into pies, jams, sauces etc[183]. Pectin or pectin-rich fruit should be added if the fruit is used in jams and preserves[183]. The fruit can also be dried, ground into a meal and added to flour for making bread etc[183]. The plant conveniently wraps up each fruit in its own 'paper bag' (botanically, the calyx) to protect it from pests and the elements. This calyx is toxic and should not be eaten.

Medicinal Uses

Disclaimer

Antitumor Diuretic Poultice.

The seed is considered to be beneficial in the treatment of difficult urination, fever, inflammation and various urinary disorders[222].
A tea made from the leaves is used in the treatment of headaches and as a wash for burns and scalds[222].
A poultice of the leaves and roots is applied to wounds[222]. An infusion of the leaves and roots is used as a wash on scalds, burns and VD sores[257].
Compounds in the plant are being investigated for antitumor activity[222].

Other Uses

None known

Cultivation details

Succeeds in any well-drained soil in full sun or light shade[200].
A polymorphic species[43].

Propagation

Seed - sow March/April in a greenhouse only just covering the seed. Germination usually takes place quickly and freely. Prick out the seedlings into individual pots of fairly rich soil when they are large enough to handle and plant them out after the last expected frosts. Consider giving them some protection such as a cloche until they are growing away well. Diurnal temperature fluctuations assist germination[170].
Division in spring[111]. Very easy, larger divisions can be planted out direct into their permanent positions. We have found that it is better to pot up the smaller divisions and grow them on in light shade in a cold frame until they are well established before planting them out in late spring or early summer.
Basal cuttings in early summer[111]. Harvest the shoots with plenty of underground stem when they are about 8 - 10cm above the ground. Pot them up into individual pots and keep them in light shade in a cold frame or greenhouse until they are rooting well. Plant them out in the summer.

Suppliers

For more details of plant suppliers please see our Suppliers Page which lists many more places to look.

Web References

[B] Data (Latin & Common names, other references) from the BONAP's Synonymized Checklist of the Vascular Flora of the United States, Puerto Rico, and the Virgin Islands.
[G] Data (Common Names, Uses, Distribution) from the USDA/ARS NPGS's GRIN taxonomic database.
[P] Data. (uses, distribution, wetland) from the USDA'a Plants database.
[HP] Links, Photos, Suppliers from Hortiplex Plant Database

References for Physalis heterophylla var. heterophylla (a possible synonym).

[B] Data (Latin & Common names, other references) from the BONAP's Synonymized Checklist of the Vascular Flora of the United States, Puerto Rico, and the Virgin Islands.
[P] Data. (uses, distribution, wetland) from the USDA'a Plants database.

References for the family Solanaceae.

[CHINA] Data Very detailed botanical/identification info. from the Flora of China.

See the PFAF Links Pages for other sources.

Also try Photos and info from the The Plants Database which has 14,000 images.

References

[K] Ken Fern
Notes from observations, tasting etc at Plants For A Future and on field trips.

[19] Stary. F. Poisonous Plants. Hamlyn 1983 ISBN 0-600-35666-3
Not very comprehensive, but easy reading.

[43] Fernald. M. L. Gray's Manual of Botany. American Book Co. 1950
A bit dated but good and concise flora of the eastern part of N. America.

[46] Uphof. J. C. Th. Dictionary of Economic Plants. Weinheim 1959
An excellent and very comprehensive guide but it only gives very short descriptions of the uses without any details of how to utilize the plants. Not for the casual reader.

[65] Frohne. D. and Pfänder. J. A Colour Atlas of Poisonous Plants. Wolfe 1984 ISBN 0723408394
Brilliant. Goes into technical details but in a very readable way. The best work on the subject that I've come across so far.

[105] Tanaka. T. Tanaka's Cyclopaedia of Edible Plants of the World. Keigaku Publishing 1976
The most comprehensive guide to edible plants I've come across. Only the briefest entry for each species, though, and some of the entries are more than a little dubious. Not for the casual reader.

[111] Sanders. T. W. Popular Hardy Perennials. Collingridge 1926
A fairly wide range of perennial plants that can be grown in Britain and how to grow them.

[161] Yanovsky. E. Food Plants of the N. American Indians. Publication no. 237. U.S. Depf of Agriculture.
A comprehensive but very terse guide. Not for the casual reader.

[170] Dremann. C. G. Ground Cherries, Husk Tomatoes and Tomatilloes. Redwood City Seed Co 1985 ISBN 0-933421-03-6
Only a small booklet but it covers the various species in some depth.

[183] Facciola. S. Cornucopia - A Source Book of Edible Plants. Kampong Publications 1990 ISBN 0-9628087-0-9
Excellent. Contains a very wide range of conventional and unconventional food plants (including tropical) and where they can be obtained (mainly N. American nurseries but also research institutes and a lot of other nurseries from around the world.

[200] Huxley. A. The New RHS Dictionary of Gardening. 1992. MacMillan Press 1992 ISBN 0-333-47494-5
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.

[222] Foster. S. & Duke. J. A. A Field Guide to Medicinal Plants. Eastern and Central N. America. Houghton Mifflin Co. 1990 ISBN 0395467225
A concise book dealing with almost 500 species. A line drawing of each plant is included plus colour photographs of about 100 species. Very good as a field guide, it only gives brief details about the plants medicinal properties.

[235] Britton. N. L. Brown. A. An Illustrated Flora of the Northern United States and Canada Dover Publications. New York. 1970 ISBN 0-486-22642-5
Reprint of a 1913 Flora, but still a very useful book.

[257] Moerman. D. Native American Ethnobotany Timber Press. Oregon. 1998 ISBN 0-88192-453-9
Very comprehensive but terse guide to the native uses of plants. Excellent bibliography, fully referenced to each plant, giving a pathway to further information. Not for the casual reader.

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Plant information taken from the Plants For A Future - Species Database. Copyright (c) 1997-2003.
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